When Moletown experiences a heatwave, Moley and Dotty hope Mishmosh can invent something to cool things down. Unfortunately, Mishmosh has "inventors block" so Moley and Dotty must help him get back his inventing mojo.

Moley must learn to Molerskate for the Great Mole Roll skating competition and discovers his parents have a hidden talent.

When an egg rolls into Moletown and a duckling hatches, Moley must play momma to the baby duck until they can find its home.

When Moley helps take care of the worms at Dotty’s Adopt a Worm Rescue Center, the Gardener sends Squirm to pose as a homeless worm and help him steal Manny.

Moley vows to not eat worms for a few days in support of Dotty’s veganism and his worm craving temptations get out of control.

When Moley spends the day with a young mole named Cheese who wants to be just like him, Moley helps Cheese recognize his own talents and uniqueness.

Mishmosh builds a tunnel digger, upsetting some that it will replace moles’ need to dig themselves. Dotty accidently gets locked in the machine and it gets activated!